photographed in Nature in different regions,
mostly in Siberia

Any use of images of this site must be accompanied with a reference to the author

All the pictures were taken in pure natural conditions without any restriction to a dragonfly freedom.

(Click an image to enlarge and see a detailed legend, often a story why this dragonfly or its very shot is interesting)


                                                            The copula right is intergeneric!!!

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last updated in February 2005


CALOPTERYGIDE (169 pictures) Last updated in March 2014

EUPHAEIDAE (46 pictures) Last updated in January 2016

CHLOROCYPHIDAE (74 pictures) Last updated in January 2016

MEGAOPODAGRIONIDAE (48 pictures) Last updated in April 2012

LESTIDAE (74 pictures) Last updated in January 2016

COENAGRIONIDAE (312 pictures) Last updated in January 2016

PLATYCNEMIDIDAE (131 pictures) Last updated in January 2016

PLATYSTICTIDAE (10 pictures of 2 species) Last updated March 2014

PROTONEURIDAE (36 pictures) Last updated in October 2015

AESHNIDAE (152 pictures) Last updated in January 2013


CORDULEGASTERIDAE (4 pictures of one species) Last updated December 2006


GOMPHIDAE (186 pictures) Last updated in January 2016

CORDULIIDAE s. l. (76 pictures) Last updated in April 2012


LIBELLULIDAE (749 pictures) Last updated in January 2016

By regions

RUSSIA and KAZAKHSTAN (661 pictures) Last updated in January 2016


JAPAN and SOUTH KOREA (124 pictures) Last updated in September 2007


GERMANY and FRANCE (79 pictures) Last updated February 2005


THAILAND (509 pictures) Last updated in November 2010


CAMBODIA (347 pictures) Last updated in January 2016


INDONESIA (113 pictures) Last updated in July 2014


ETHIOPIA (96 pictures) Last updated in January 2013


Miscelaeous fragments of Russian odonatological literature translated by O. Kosterin into English

View the list of dragonfly taxa preserved in Siberian Zoological Museum at the Institute of Systematicsand Ecology of Animals. Novosibirsk.
(including images of Belyshev's type specimens)

By clicking highlighted references you'll see the very text (without illustrations)

Zaika, V.V., Kosterin, O.E. 1987. [Community structure and ethological aspects of dragonflies of Lake Manzherok (Altai)]. In: [Ecology and Geography of Arthropods of Siberia]. Nauka (Siberian Division), Novosibirsk. pp. 73-74 (in Russian)

Kosterin, O.E. 1987. Discovery of East-Asiatic dragonfly (Odonata, Libellulidae) at the Manzherock Lake (Altay). In: [Insects, Mites, and Helmints. New and little-known species of the fauna of Siberia]. Nauka (Siberian Division), Novosibirsk. pp. 57-63. (in Russian)

Kosterin, O.E. 1987. [Dragonfly community of Lake Manzherok]. In: [Problems of formation of communities terrestrial and aquatic biocenoses]. OmGPI, Omsk. pp. 76-92. (in Russian)

Kosterin O.E. 1989. To the dragonfly fauna of Altai Mts. Theses of the All- Union Scientific-Practical Conference "Problems of Ecology of Mountainous Regions" 9-13 October 1989. Dushanbe. pp. 6-10 (in Russian)

Kosterin, O.E. 1992. New distribution records of Somatochlora sahlbergi Trybom (Odonata, Corduliidae). Acta Hydroentomo- logica Latvica - Vol. 2 - P. 22-26.Russian text is available here

Zaika, V.V., O.E.Kosterin. 1992. Some interesting observations of dragonflies (Odonata) in South Tuva. Acta Hydroentomologica Latvica Vol. 2 - P. 81-84.Russian text is available here

Kosterin, O. E., V. V. Zaika. Materialy oi strekozam (Odonata, Insecta) k Krasnoy Knige Respubliki Altai. [Materials on dragonflies (Odonata, Insecta) Diurna) for the Red Book of Altai Republic]. In: Materialy k Krasnoi knige Respubliki Altai (zhivotnye). [Materials for the Red Data Book of Altai Republic (animals)], N. P. Malkov, ed. - Gorno-Altaisk State University - Gorno-Altaisk - 1995 - p. 96-97 (in Russian)

Kosterin, O. 1996. Dragonflies (Odonata) of the city of Omsk. Acta Hydroentomologica Latvica - Vol. 3 - P. 10-21.

Kosterin, O.E. 1996. Items: Sympetrum croceolum Selye 1883; Ischnura pumilio Charpentier, 1825; Nechalenia speciosa Charpentier, 1840. Zaika, V.V., Kosterin O.E. Item: Anax parthenope Selys, 1839. In: The Red Book of Altai Republic. Animals. Novosibirsk. pp. 44-49, 54-56. (in Russian)

Kosterin, O. E. 1998. Vazhnye nakhodki v faune strekoz (Odonata) Daurskogo zapovednika i ego okrestnostei. [Important findings in the dragonfly (Odonata) fauna of the Dahurian Nature Reserve and its environs. In: Problems of Entomology in Russian]. Abstracts for XI Congress of Russian Entomological Society, Sanct-Petersburg, - Vol. 1 - P. 210-211< (in Russian). Available in Russian and English)

O.E. Kosterin. 1999. Dragonflies (Odonata) of the Daurskii State Nature Reserve and its surroundings. In: Nasekomye Daurii i sopredel'nykh territorii (Sbornik nauchnykh trudov). Vypusk II. [Insects of Dauria and Adjacent Terrtories. (Collected scientific works). Issue II]. - Novosibirsk - P. 5-40. (in Russian, Engl. res.)
English translation available here

O. E. Kosterin. 2000. Observation of an intergeneric copulation between a male Cordulia aenea (l.) and a female Epitheca bimaculata (Charp.) (Anisoptera: Corduliidae). Notulae Odonatologicae, Vol. 5, No. 5, p. 55-56.

Krasnaya Kniga Chitinskoi oblasti i Aginskogo Buryatskogo Avtonomnogo okruga. Zhivotnye [Red Book of Chita Province and Aginskii Buryat Autonomous Region. Animals.]. A. M. Vozmilov, ed. - 2000 - Poisk - Chita. Items: by O. E. Kosterin: Strelka v-chernoe, Cercion v-nigrum (Needham, 1930) - p. 140-141; Dedka Maaka, Anisogomphus maackii (Selys, 1872) [= Gomphus m-flavum (Selys, 1872)] - p. 141-142; Dozorshchik iyul'skii, Anax parthenope Selys, 1839, ssp. julius Brauer, 1866 - p. 142-143; Brodyazhka ryzhaya, Pantala flavescens (Fabricius, 1798) - p. 145-146. By O. E. Kosterin and E. A. Maksimenko: Babka Zal'berga - Somatochlora sahlbergi Trybom, 1889 - p. 143-144. (in Russian)

Kosterin, O. E., Haritonov, A. Y., Inoue, K. 2001. Dragonflies of the part of Novosibirsk Province east of the Ob', Russia. - Sympetrum Hyogo - Vol. 7/8 - p. 24-49 (in English and Japanese).

Kosterin, O. E., Zaika, V. V. 2001. Odonate fauna of the Tyva Republic, or Tuva (Central Siberia). - The Fifteenth International Symposium of Odonatology. Abstracts. Novosibirsk, Russia, July 9-19, 2001 - pp. 10-11.

Krasnaya kniga Respubliki Tyva. Zhivotnye [Red Data Book of Tyva Republic. Animals, N. I. Putuintsev, ed.]. 2002. SO RAN Press, Geo Branch, Novosibisrk, 167 p. Articles: Kosterin, O. E. Dedka shiporogii Ophiogomphus spinicornis Selys, 1878 - p. 14. (in Russian)

Kosterin, O. E. 2002. Western borders and isolates of some generally East Asian odonate species in South Siberia and their putative origin. - In: The first Symposium of the SIOROEA (S.I.O. Regional Office in East Asia. Abstracts and program. - National Science Museum, Daejeon, July 26-29 2002 - p. 12-13.

Kosterin, O. E., V. V. Zaika. 2003. Odonatological expeditions to the Tyva Republic (Tuva) 2000-2002. IDF-Report 5: 1-32.

Kosterin, O. E. 2004. Odonata of the Daurskii State Nature Reserve area, Transbaikalia, Russia. Odonatologica 33 (1): 41-71.

Kosterin, O. E. 2004. Some Odonata collected in Aldan Ulus of Sakha (Yakutia) Republic in Late June 2002. Notulae Odonatologicae 6 (3): 27-31.

Kosterin, O. E., E. I. Malikova, O. E. Haritonov. 2004. Critical species of Odonata in the Asian part of the former USSR and the Republic of Mongolia. International Journal of Odonatology 7: 341-370.

Dubatolov, V. V., R. Yu. Dudko, V. G. Mordkovich, O. B. Korsun, S. E. Chernyshev, D. V. Logunov, Yu. M. Marusik, A. A. Legalov, S. V. Vasilenko, L. G. Grishina, G. S. Zolotarenko, A. V. Barkalov, V. P. Petrova, P. Ya. Ustjuzhanin, S. Yu. Gordeev, V. K. Zihcnenko, M. G. Ponomarenko, I. I. Lyubechanskii, N. N. Vinokurov, O. E. Kosterin, E. I. Malikova, A. L. L'vovskii, E. A. Maksimenko, E. E. Malkov, A. N. Streltzov, S. G. Rudykh, D. A. Mil'ko. 2004. Biodiversity of the Sokhondo Nature Reserve. Arthropoda. Sokhondo Nature Reserve, Institute of Systematics and ecology of Animals SB RAS, Novosibirsk, Chita, 415 p. (in Russian) [a part on Odonata, pp. 81-87, by O. E. Kosterin, E. I. Malikova, E. A. Maksimenko, O. V. Korsun].

Dumont, H., A. Y. Haritonov, O. E. Kosterin, E. I. Malikova, O. G. Popova. 2005. A review of the Odonata of Kamchatka peninsula, Russia. Odonatologica 34 (2): 131-153

Kosterin, O. E. 2005. Western range limits and isolates of eastern odonate species in Siberia and their putative origins. Odonatologica 34 (3): 219-242.

Kosterin, O. E., Dubatolov V. V. 2005. A dragonfly (Odonata) collection from the Bol'shekhekhtsirskii State Nature Reserve (Khabarovskii Krai, Russia) - Fauna of Russian Far East: collected research papers. Volume 5 - Blagoveshchensk State Pedagogical University, Blagoveshchensk - P. 9-14.

Kosterin O. R., Vikhrev N. E. 2006. Odonata seen during three days in a populated lowland part of Cambodia. Malangpo 21: 212-217.

Kosterin, O. E., O. V. Korsun. 2006. A collection of Odonata from the Argun' (Hailar) River basin in Transbaikalia, East Siberia, Russia. - Notulae Odonotologicae 6: 81-85.

Bernad, R., O. E. Kosterin. 2007. Odonatologiczne impresje z Riowniny Wasjuganu, zachodnia Syberia. Odonatrix 3 (2): 50-58.

Kosterin, O. E. 2007. The first record of Anax of the West Siberian Plain: A. p. parthenope Selys in Omsk (Anisoptera: Aeshnidae). Notulae Odonatologicae 6 (10): 112-115.

Malikova E. I., O. E. Kosterin, V. V. Dubatolov. 2007. A dragonfly (Odonata) collection from the Bolshekhekhtsirskii State Nature Reserve (Khabarovskii Krai, Russia). II. Seasons 2006 and 2007. In: Zhivotnyi mir Dal'nego Vostoka [Animal World of the Far East], A. I. Streltzov ed., Issue 6, p. 5-9. A PDF-file

Bernard, R., O. Kosterin. 2008. Field notes of two hunters for Nechalennia speciosa in boggy Vasyugan Plain, West Siberia. IDF-Report 12: 1-34. A PDF-file

Kosterin, O. E. 2007. Strekozy (Insecta, Odonata) Akademgorodka [Dragonflies (Insecta, Odonata) of Akademgorodok]. In: Priroda Akademgorodka: 50 let spustya [The Nature of Akademgorodok: 50 years passed], I. F. Zhimulev (ed.), SO RAN Publishing House, Novosibirsk, pp. 74-91 (In Russian).

Kosterin, O. E., N. E. Vikhrev. 2008. Odonatological field notes of two January trips to south-eastern Thailand in 2005 and 2006. Malangpo 22: 222-236.

Kosterin, O. E. 2008. Observations on the crepuscular flight in Aeshna viridis Eversman in Omsk Province, West Siberia (Anisoptera: Aeshnidae). Notulae odonotalogicae 7 (2): 18-20.

Kosterin, O. E., L. V. Sivtseva. 2009. Odonata of Yakutia (Russia) with description of Calopteryx splendens njuja ssp. nov. (Zygoptera: Calopterygidae). Odonatologica 38 (2): 93-202.

Kosterin, O. E., I. I. Lyubechanskii. 2009. Odonata collection from the Bureinskii State Nature Reserve, Khabarovskii Krai, Russia. Notulae odonatologicae 7 (3): 25-27.

Kosterin, O., N. Vikhrev. 2009. Some new provincial records of Odonata made in Thailand in 2007-2009 and a new record for Vietnam. Agrion 13 (2): 75-79, only this paper.

Kosterin, O. E. 2009. Observation of flying activity of Ophiogomphus spinicornis Selys, 1978 (Insecta, Odonata. Gomphidae) in Shiveelig-Khem River plain, Ubsunur Hollow, Southern Tuva. In: Nature of Ubsu-Nur Hollow Reserve. Transaction of "Ubsunur Hollow" State Biospheric Reserve. Issue 1, Darma Press, Krasnoyarsk, p. 226-232. (in Russian)

Bernard, B., O. E. Kosterin. 2010. Biogeographical and ecological description of Odonata of eastern Vasyugan Plain, West Siberia, Russia. Odonatologica 39: 1-28.

Kosterin, O. E., P. G. Gorbunov. 2010. Notes on the Odonata fauna of Kazakhstan, including the first record of Ischnura evansi Morton (Zygoptera: Coenagrionidae). // Notulae odonatologicae 7: 45-48.

Kosterin, O. E. 2010. Siberian taxonomical problems concerning European odonate species. 1st European Congress on Odonatology, Programme and abstracts, 2-5 July, Vaira~o - Vila do Conde, Portugal, p. 28

Kosterin, O.E., S.N. Borisov. 2010. Dragonflies (Odonata) of the Dzhungarskiy Alatau mountains, South-East Kazakhstan. Eurasian Entomological Journal 9: 299-302.

Kosterin, O.E., V.V. Zaika. 2010. Odonata of Tuva, Russia. International Journal of Odonatloogy 13: 277-328.

Kosterin, O.E. 2010. A glance at the Odonata of the Cambodian coastal regions: end of dry season in 2010. International Dragonfly Fund Report 29: 1-75.

Kosterin, O., N. Vikhrev, V. Zinchenko, A. Korshunov. 2011. More Odonata records from Thailand. Agrion 15: 12-16.

Kosterin, O.E., N.V. Skalon, T.N. Skalon. 2011. Interesting findings of Odonata in the Kuzhentskiy Alatau Mts. north-eastern foothills. Amurian Zoological Journal 3: 124-127

Kosterin, O. E. 2011. Odonata of the Cambodian coastal regions revisited: beginning of dry season 2010. International Dragonfly Fund Report 40: 1-108.

Kosterin, O.E., V.V. Zaika. 2011. Fauna of dragonflies and damselflies (Odonata) of Tuva. // Amurian Zoological Journal 3: 210-245 (in Russian)

Kosterin, O.E., J. Holden. 2011. Some photographic records of Odonata in Cambodia. International Dragonfly Fund Report 42: 1-6.

Day, L., D. Farrell, A. Guenther, M. Hamalainen, E. Klimsa, A. Korshunov, O. Kosterin, N. Makbun, A. Pelegrin, U. Roeder, R. Ruangrong, N. Vikhrev. 2012. New provincial records of Odonata from Thailand mostly based on photographs. Agrion 16: 16-24.

Kosterin O. 2012. Odonata of the Cambodian coastal regions in late rainy season of 2011. // International Dragonfly Fund Report, Vol. 45, pp. 1-102.

Kosterin O.E. 2012. A rapid survey of Odonata on Bokor Plateau, Preah Monivong National Park, Cambodia. // Cambodian Journal of Natural History, Vol. 2012, pp. 75-86.

Kosterin, O.E., N. Makbun, P. Dawwrueng. 2012. Burmagomphus asahinai sp. nov., a new species from Cambodia and Thailand, with a description of the male of B. gratiosus Chaoi, 1954. // International Journal of Odonatology, Vol. 15, pp. 275-292.

Kosterin, O.E., G. Chartier, J. Holden, F.S. Mey. 2012. New records of Odonata from Cambodia, based mostly on photographs. Cambodian Journal of Natural History 2012: 150-163.

Skalon, N.V. (ed.). 2012. Krasnaya Kinga Kemerovskoi oblasti. T. 2. Redkie i nakhodyashchiesya pod ugrozoi ischeznoveniya vidy zhivotnykh. [Red Data Book of Kemerovo Province. V. 2. Rare and threatened animal species] - Asia-Print, Kemerovo, 192 p. Articles: Dronzikova, M.V., O.E. Kosterin, T.N. Skalon: Calopteryx japonica Selys, 1869 - p. 23; Anax parthenope (Selys, 1839) - p. 24; Skalon, T.N., O.E. Kosterin, N.V. Skalon:. Gomphus vulgatissimus (Linnaeus, 1758) - p. 26 (in Russian).

Kosterin O.E. 2013. Brief Odonata survey in North Ethiopia during heavy rainy season of 2012. // International Dragonfly Fund Report, Vol. 56, pp. 1-54.

Medvedev, A.F.m O.E. Kosterin, E.I. Malikova, W. Schneider. 2013. Description date of Somatochlora exuberata Bartenev, Leucorrhinia intermedia Bartenev and Sympetrum vulgatum grandis Bartenev, the fate of A.N. Bartenev’s type specimens and designation of the lectotype of L. intermedia (Anisoptera: Corduliidae, Libellulidae). Odonatologica – Vol. 42 – pp. 211-228.

Kosterin O.E. 2013. Somatochlora arctica (Zett.) seems to migrate to escape wildfires (Anisoptera, Corduliidae). Notulae odonatologicae – Vol. 8 – pp. 35-36.

Popova. O.N., S.N. Borisov, O.E. Kosterin, B. Kiauta. 2013. Obituary. Anatoly Yuryevich Haritonov. Odonatologica – Vol. 42 – pp. 409-425.

Kosterin, O.E. 2014. Odonata of the sourth-west and north-east of Cambodia as studied in early rainy season of 2013. International Dragonfly Fund Report 67: 1-94.

Kosterin, O.E. 2014. Notes on infraspecific variation of some Gomphidae (Odonata) species in Cambodia. Internaional Dragonfly Fund Report 68: 1-16.

Kosterin, O.E. 2014. Odonata briefly observed on the islands of Bali and Lombok, Lesser Sundas, Indonesia, in the late Febriary 2014. International Dragonfly Fund Report 74: 1-48.

Kosterin, O.E., G. Chartier. 2014. Two more Odonata species recorded from Cambodia. Cambodian Journal of Natural History 2014: 8-11

Kosterin OE, Constant J, Wilson KDP. 2014. Neotype of Pseudagrion approximans Selys, 1867 designated to resolve a nomenclatorial confusion in the genus Aciagrion Selys, 1891 (Odonata: Coenagrionidae) // International Journal of Odonatology 17: 161-172. DOI:10.1080/13887890.2014.959075

Borisov, S.N., Kosterin O.E. 2014. On the fauna of Odonata of Chukotka and other northern regions of the Holarctic. // Eurasian Entomological Journal. 13 (4): 315-320.

Borisov, S.N., Kosterin O.E. 2014. Dragonflies and damselflies (Odonata) of north-eastern Kazakhstan. // 13 (4): 339-345.

Kosterin O.E. 2015. Taxonomical notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 1. Indolestes gracilis expressior ssp. nov. from eastern Cambodia. // International Dragonfly Fund – Report 81: 1-11.

Kosterin O.E., R. Poggi. 2015. Taxonomical notes on Indolestes Fraser, 1922 (Lestidae, Zygoptera). 2. Indolestes birmanus (Selys, 1891) is bona species. // International Dragonfly Fund – Report 81: 13-20.

Kosterin O.E., H. Karube, R. Futahashi. 2015. Two new subspecies of Hemicordulia tenera Lieftinck, 1930 (Corduliidae) from Cambodia and Thailand. // International Dragonfly Fund – Report – Vol. 82 – P. 1-19.

Kosterin O.E. Risiophlebia guentheri sp. nov. (Odonata, Libellulidae) from southeastern Indochina. // Zootaxa 3964: 138-145.

Kosterin O.E. 2015. Taxonomic and faunal notes on Macromia Rambur, 1842 from Cambodia (Odonata: Macromiidae). // Odonatologica 44: 117-151

217. Kosterin O.E. 2015. Onychargia priydak sp. nov. (Odonata, Platycnemididae) from eastern Cambodia. // International Journal of Odonatology. Vol. 18. Iss. 2. P. 157-168.

Zhang H.-m., Kosterin O.E., Cai C.-h. 2015. New species and records of Burmagomphus Williamson, 1907 (Odonata, Gomphidae) from China. // Zootaxa. Vol. 3999. Issue 1. P. 62-78

Kosterin O.E. 2015. Prodasineura hoffmanni sp. nov. (Odonata, Platycnemididae, Disparoneurinae) from eastern Cambodia. // Zootaxa. Vol. 4027. Iss. 4. P. 565-577.

Kosterin O.E. 2015. Ischnura foylei sp. nov. (Odonata, Coenagrionidae) from the highlands of Sumatra. // Zootaxa. Vol. 4032. Iss. 4. P. 179-189.

Kosterin O.E. 2015. On the Odonata of North Kazakhstan Province. I. First data on Petropavlovsk. // International Dragonfly Fund Report. Vol. 86. P. 1-26.

Kosterin O.E. 2015. Odonata registered on a short excursion to Kyshtovka District, Novosibirsk Province. // International Dragonfly Fund Report. Vol. 86. P. 29-46.

Kosterin O.E. 2015. Dry season Odonata of the Cardamonean coast (Cambodia and Thailand) revisited. // International Dragonfly Fund Report. Vol. 89. P. 1-36.

Kosterin, O.E. & Yokoi N. 2016. Asiagomphus reinhardti sp. nov. (Odonata, Gomphidae) from eastern Cambodia and southern Laos. // Zootaxa Vol. 4103. Issue 1. P. 35-42. Kosterin, O.E. 2016. Microgomphus alani (Odonata, Gomphidae) sp. nov. from Cambodia. // Zootaxa Vol. 4114. Issue 3. P. 341-350.
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