Clossiana titania (Esper, 1793) (= Boloria titania).

Range: boreal forests of the temperate Eurasia.

ssp: staudingeri (Wnukowsky, 1929); South and Central Siberia.

Wings open.

A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999. Photo: O. Kosterin.